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"Lolicornes", c'est un monde joyeux et coloré peuplé de créatures mythiques ou éteintes dans notre monde : licornes, cyclopes, dodos... Mais derrière cette façade se cache en fait une société hautement inégalitaire : dès son plus jeune âge, chaque habitant de Loliland est rangé dans la catégorie "beau" ou "moche" en fonction d'un certain nombre de critères très subjectifs. La vie des "beaux" est idyllique alors que celle des "moches" est remplie de corvées et de dangers. Mais l'équilibre de cette société va être bouleversé par l'arrivée de Penny, une gamine humaine amnésique. Plus grave encore : une crise de "grise-mine" va attaquer une bonne partie de la population... mais uniquement les beaux ! Mamy Loly, dirigeante implacable de Loliland, va directement lier l'arrivée de Penny à cette maladie et charger Moe, un moche à tout faire, et ses copains pas beaux d'emmener Penny très loin... et de s'en débarrasser !