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Elder Tale est le jeu en ligne qui rassemble le plus grand nombre de joueurs dans un monde où la nature a repris ses droits. Mais lors de la sortie de sa douzième extension, tous ceux qui s’étaient connectés se retrouvent prisonniers du jeu. Ignorant comment ils sont arrivés là et s’ils pourront en sortir un jour, beaucoup perdent la volonté d’avancer. Mais Shiroe, un des joueurs les plus connus et les plus respectés du jeu refuse d’abandonner. Aidé de ses compagnons Akatsuki et Naotsugu, il va tenter de convaincre les « Aventuriers » de ne pas se laisser abattre et de bâtir une nouvelle société dans ce monde virtuel qui est maintenant le leur.
Le 15 juillet 2043, sortait le VRMMO immersif <Infinite Dendrogram> intégrant «l'Embryo», un système unique se développant en une infinité de variétés selon chaque joueur. S'appuyant sur des prouesses techniques qui étaient jusque-là inaccessibles aux VRMMO, <Infinite Dendrogram> est devenu en un clin d'œil un phénomène mondial. Mukudori Reiji débute sa vie à Tokyo après avoir réussi ses examens d'entrée à l'université et, pour célébrer la fin de sa longue période de révision, il lance <Infinite Dendrogram> pour enfin y rejoindre son frère...
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