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Tanjiro, Zenitsu, Inosuke et Nezuko séjournent dans un village pour soigner les blessures qu'ils ont subies au cours de leur dernière mission. Dans ce village, Tanjiro assiste à une cérémonie de mariage et aperçoit la mariée qui a l'âge de Nezuko. Il commence alors à imaginer l'avenir de sa soeur bien-aimée. Pendant ce temps, il apprend la légende d'une fleur qui apporterait le bonheur à ceux qui la trouvent.... Demon Slayer est le titre de tous les records. Chaque fois que le manga de Koyoharu Gotouge est cité, un nouvel exploit est atteint ! Un an après sa publication en France, le premier tome est entré dans les dix plus grosses ventes de manga en 2020 (Classement Gfk). Le film Le train de l'infini a généré 32,5 milliards de yens de recette, il a battu le record du box-office au Japon, détrônant ainsi le Voyage de Chihiro ! La chanson thème de l'anime domine aussi les charts nippons...
A notre époque, d'étranges portails sont apparus sur Terre, connectant la planète à des donjons peuplés de monstres. Pour les combattre, l'humanité peut compter sur les chasseurs, des humains ayant développé des capacités surnaturelles. Chasseur de rang E, Sung Jinwoo se rend un jour dans un donjon de bas niveau avec son équipe. Mais l'escadron se retrouve piégé dans une salle remplie de monstres surpuissants...
L'histoire débute avant la chute du monde et le règne des vampires. Glenn Ichinose, le mentor de Yuichirô, est alors âgé seulement de 16 ans. Il est sur le point d'intégrer la prestigieuse académie de magie de Shibuya 1 dirigée par la famille Hiiragi. On dit qu'il faut pourtant être le meilleur et être né dans les clans les plus prestigieux pour y parvenir. Glenn vient d'un clan mineur et rebelle, mais il est prêt à y subir les pires humiliations pour servir son ambition. Il va y croiser une jeune femme à laquelle il avait fait une promesse il y a bien longtemps... Alors que le monde court à sa perte, tous, Glenn comme les autres, sont guidés par une terrible soif de puissance...!
