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Un Japon où les humains et des esprits surnaturels, les Ayakashi, coexistent. Oni... Esprits-Renards, Chats-Spectres... Grâce à leurs pouvoirs extraordinaires et à leur beauté fascinante, les Ayakashi forment le coeur et l'élite du pays. Lorsqu'un Ayakashi identifie une jeune humaine comme sa "fiancée", elle reçoit la promesse d'un amour inconditionnel. Yuzu est une lycéenne qui vit dans l'ombre de sa soeur cadette, la fiancée d'un Esprit-Renard. Méprisée et privée d'affection par sa famille, elle endure chaque jour une profonde solitude. Un soir, elle croise dans la rue un jeune homme d'une beauté saisissante. Ce garçon aux yeux rouges n'est autre que Reiya Kiryûin, héritier de la famille la plus puissante des Ayakashi et futur maître de tous les Oni.
Osaka, de nos jours. La jeune Utsui est en terminale et est élevée dans une famille queer, dans un environnement où aimer qui on veut est plus qu’accepté. Elle-même, cependant, se débat avec son incapacité d’aimer de manière romantique. Parmi les personnes de passage dans sa maison, elle s’attache à Tamon, une personne étrange qui a pour habitude de converser avec des êtres surnaturels qu’iel seul·e peut voir. En réponse à l’appel d’Utsui, un mystérieux petit être ami de Tamon apparaît… mais il a oublié son propre nom.
Après avoir été trahie par sa seule amie et son propre fiancé, Mayumi décide de commencer une nouvelle vie à la campagne, où elle ne connaît personne, en tant que professeure au collège. À peine est-elle arrivée qu'elle se retrouve à faire un plongeon dans l'eau avec une fille aux airs de délinquante. Elle découvre ensuite qu'il s'agit de Rin, la gérante de l'endroit où elle va habiter, la villa de la Mouette. Ici, sous le même toit, débute la cohabitation de deux femmes malmenées par le destin.
