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Le Joker, connu pour sa folie, aussi meurtrière que machiavélique, se retrouve face à Batman une nouvelle fois, mais que se passerait-il si l'on inversait les rôles ? Jack Napier, alias le Joker guéri et libéré de ses pensées criminelles se présente aux élections pour devenir le nouveau maire de Gotham. De l'autre côté, le Chevalier Noir tombe dans la criminalité pour sa violence mise à jour par des réseaux sociaux. L'ancien Prince du Crime endosse le rôle du Chevalier Blanc pour sauver Gotham.
Enquêtant sur l'existence et les propriétés de différents métaux disséminés à travers la planète depuis des millénaires, Batman découvre un portail ouvrant sur un anti-multivers ; des dimensions parallèles où l'Histoire a déraillé et où des Chevaliers Noirs terrifiants ont supprimé les membres de la Ligue de Justice. Aujourd'hui, ces derniers décident d'envahir notre dimension.
Darkseid a de nombreuses fois tenté de conquérir la Terre et de réduire à néant les super-héros qui la défendent. Mais aujourd'hui, il y est parvenu. Lors d'un combat contre la Ligue de Justice, le seigneur d'Apokolyps a déchaîné toute la puissance de l'équation d'anti-vie, faisant ainsi du monde un enfer habité d'individus contaminés et hystériques qui se dévorent les uns les autres. Et face au chaos planétaire de l'anti-vie, les héros sont aussi vulnérables que désemparés.
