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Quand le C omédien, justicier au service du gouvernement, se fait défenestrer, son ancien allié, Rorschach, mène l'enquête. I l reprend rapidement contact avec d'autres héros à la retraite dont le D r Manhattan, surhomme qui a modifié le cours de l'histoire. Alors qu'une guerre nucléaire couve entre les USA et l'URSS, tous s'interrogent : qui nous gardera de nos Gardiens ?
Spider-Man se voit offrir un nouveau costume de la part de son nouveau meilleur ami... Tony Stark ! Et les liens tissés aujourd'hui vont déterminer le choix de Spidey durant le conflit à venir. Les Quatre Fantastiques, quant à eux, enquêtent sur un mystérieux objet tombé du ciel tandis que l'équipe la plus secrète de l'univers Marvel, les Illuminati, se reforme. Enfin, les New Warriors deviennent les stars d'une émission de téléréalité et parcourent les routes des États-Unis à la recherche de nouveaux exploits.
Le père de l'auteur, Vladek, juif polonais, rescapé d'Auschwitz, raconte sa vie de 1930 à 1944, date de sa déportation. Ce récit est rapporté sous la forme d'une bande dessinée dont les personnages ont une tête d'animal : les juifs sont des souris, les nazis des chats, les Polonais des porcs et les Américains des chiens.
