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Le Joker, connu pour sa folie, aussi meurtrière que machiavélique, se retrouve face à Batman une nouvelle fois, mais que se passerait-il si l'on inversait les rôles ? Jack Napier, alias le Joker guéri et libéré de ses pensées criminelles se présente aux élections pour devenir le nouveau maire de Gotham. De l'autre côté, le Chevalier Noir tombe dans la criminalité pour sa violence mise à jour par des réseaux sociaux. L'ancien Prince du Crime endosse le rôle du Chevalier Blanc pour sauver Gotham.
Quel lien secret Batman et le Joker partagent-ils avec une jeune fille mystérieuse ? Kidnappée par le Joker, le Chevalier noir doit plonger dans les profondeurs de Gotham City et s'engager dans une course contre la montre pour la retrouver. Les enjeux sont importants, et pour Batman, c'est personnel !
Spider-Man se voit offrir un nouveau costume de la part de son nouveau meilleur ami... Tony Stark ! Et les liens tissés aujourd'hui vont déterminer le choix de Spidey durant le conflit à venir. Les Quatre Fantastiques, quant à eux, enquêtent sur un mystérieux objet tombé du ciel tandis que l'équipe la plus secrète de l'univers Marvel, les Illuminati, se reforme. Enfin, les New Warriors deviennent les stars d'une émission de téléréalité et parcourent les routes des États-Unis à la recherche de nouveaux exploits.
