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Printemps 1964, à Naples, en Italie. Lucia, dix ans, vit au sein d'une troupe de cirque avec son père, Roberto, et affectionne particulièrement son lion, Cyrus. Mais après un terrible incendie qui a tout ravagé, elle décide de surmonter sa tristesse et d'échafauder un plan... Des lions, il y en a en Afrique du sud ! Jamais à court d'idée, elle fugue et embarque à bord d'un cargo, avant d'être rejointe in-extremis par son père. À leur arrivée à Durban, ils découvrent un pays en proie aux inégalités. Happés par une manifestation malgré eux, Roberto qui tente de défendre Nelson, un jeune garçon noir battu par un policier sans scrupules, se fait arrêter... Lulu et Nelson que tout tend à séparer se retrouvent unis autour d'un même combat : la quête de la liberté.
En pénétrant dans la luxuriante forêt multicolore du Royaume de Pluie, les Dragonnets du Destin espéraient trouver un répit - les lieux ont la réputation d'être pacifiques autant que splendides. Mais rien ne se passe comme prévu : menace d'une mystérieuse créature, disparition de dragons... Gloria décide de passer à l'action. Accompagnée de ses amis, elle s'enfonce dans la jungle et se retrouve en plein territoire ennemi !
