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Chargés par la division spéciale de la police métropolitaine de retrouver le concepteur du système Xenonine, Shôta et Rokurô se font attaquer par un mystérieux adversaire, équipé, lui aussi, d'une armure de combat. Ils parviennent à le repousser, avec l'aide d'un voyou du quartier surnommé Shôgun, mais l'ennemi ne tarde pas à réapparaître et à semer le chaos en ville. Shôta pourra-t-il l'arrêter à temps ?
Un Japon où les humains et des esprits surnaturels, les Ayakashi, coexistent. Oni... Esprits-Renards, Chats-Spectres... Grâce à leurs pouvoirs extraordinaires et à leur beauté fascinante, les Ayakashi forment le coeur et l'élite du pays. Lorsqu'un Ayakashi identifie une jeune humaine comme sa "fiancée", elle reçoit la promesse d'un amour inconditionnel. Yuzu est une lycéenne qui vit dans l'ombre de sa soeur cadette, la fiancée d'un Esprit-Renard. Méprisée et privée d'affection par sa famille, elle endure chaque jour une profonde solitude. Un soir, elle croise dans la rue un jeune homme d'une beauté saisissante. Ce garçon aux yeux rouges n'est autre que Reiya Kiryûin, héritier de la famille la plus puissante des Ayakashi et futur maître de tous les Oni.
Jonathan April rêve de devenir journaliste, comme son grand frère. Lorsque ce dernier lui demande de le remplacer pour une interview, il accepte sans se douter qu'il rencontrera le célèbre tueur en série " Slaughter Kister ". Après avoir malencontreusement attiré son attention, sa vie bascule...
