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Toujours sur les traces de l’armée de Neptune et de Lyssa, Icare et Kalio mettent le cap au nord, vers l’île aux quatre couleurs. Ils arrivent en pleine course annuelle : les quartiers s’affrontent en quête du pouvoir.
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Pour les vacances, Migali accueille son ami Gredin chez elle. Génial, non? Pas si sûr pour le prince grenouille qui doit s'adapter au rythme endiablé de la famille araignée. Mais la rentrée est déjà là, l'occasion de retrouver tous les copains et de faire la connaissance d'Orson, ce mystérieux prince venu du Royaume des neiges.
