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Le Joker, connu pour sa folie, aussi meurtrière que machiavélique, se retrouve face à Batman une nouvelle fois, mais que se passerait-il si l'on inversait les rôles ? Jack Napier, alias le Joker guéri et libéré de ses pensées criminelles se présente aux élections pour devenir le nouveau maire de Gotham. De l'autre côté, le Chevalier Noir tombe dans la criminalité pour sa violence mise à jour par des réseaux sociaux. L'ancien Prince du Crime endosse le rôle du Chevalier Blanc pour sauver Gotham.
Des années plus tôt, Batman subit sa plus terrible défaite lorsque le Joker tua le second Robin, Jason Todd, à coups de barre à mine. Le Chevalier Noir a néanmoins réussi à poursuivre sa mission, épaulé par son précédent acolyte, Dick Grayson. Mais ce souvenir douloureux est ravivé lorsqu'un nouveau justicier, aux moyens expéditifs, débarque à Gotham. Instaurant l'effroi dans les rangs de la pègre, il utilise de plus l'ancien alias du Joker : Red Hood.
Après avoir enlevé le Psycho-Pirate de la prison de Santa Prisca, Batman apparaît plus que jamais comme l'homme à abattre pour son ennemi juré, Bane. Bien décidé à briser le héros une fois pour toutes, Bane l'attaque sur son point faible : sa famille recomposée, Nightwing, Red Hood et Robin !
