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Qui a tué le Gardien, cet être mystérieux qui observe le monde depuis des millénaires ? Le Docteur Strange, la Panthère Noire, Emma Frost, le Punisher, Ant-Man et bien d'autres héros vont alors mener l'enquête afin d'identifier le meurtrier. Mais au cours de leurs investigations, de nombreux secrets sont révélés...
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