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Le Joker, connu pour sa folie, aussi meurtrière que machiavélique, se retrouve face à Batman une nouvelle fois, mais que se passerait-il si l'on inversait les rôles ? Jack Napier, alias le Joker guéri et libéré de ses pensées criminelles se présente aux élections pour devenir le nouveau maire de Gotham. De l'autre côté, le Chevalier Noir tombe dans la criminalité pour sa violence mise à jour par des réseaux sociaux. L'ancien Prince du Crime endosse le rôle du Chevalier Blanc pour sauver Gotham.
Darkseid a de nombreuses fois tenté de conquérir la Terre et de réduire à néant les super-héros qui la défendent. Mais aujourd'hui, il y est parvenu. Lors d'un combat contre la Ligue de Justice, le seigneur d'Apokolyps a déchaîné toute la puissance de l'équation d'anti-vie, faisant ainsi du monde un enfer habité d'individus contaminés et hystériques qui se dévorent les uns les autres. Et face au chaos planétaire de l'anti-vie, les héros sont aussi vulnérables que désemparés.
Depuis la bataille de Geonosis qui concluait l'Épisode II, les forces séparatistes, dirigées par le Comte Dooku, tentent de faire sécession. Pour les empêcher de détruire une fabrique de Clones, un commando de Jedi - parmi lesquels Obi-Wan Kenobi et Anakin Skywalker - est envoyé sur la planète aquatique Kamino. Ils découvrent que Jango Fett, bien que mort, joue encore un rôle important dans cette guerre.
