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Le Joker, connu pour sa folie, aussi meurtrière que machiavélique, se retrouve face à Batman une nouvelle fois, mais que se passerait-il si l'on inversait les rôles ? Jack Napier, alias le Joker guéri et libéré de ses pensées criminelles se présente aux élections pour devenir le nouveau maire de Gotham. De l'autre côté, le Chevalier Noir tombe dans la criminalité pour sa violence mise à jour par des réseaux sociaux. L'ancien Prince du Crime endosse le rôle du Chevalier Blanc pour sauver Gotham.
Quand il avait six ans, Bruce Wayne a vu ses parents se faire assassiner sous ses yeux. Après un entrainement intensif, il revient à Gotham City pour mener une guerre sans merci contre le crime... mais ce ne sera pas facile. Face à la corruption des autorités de la ville et leurs liens avec la pègre, Bruce, sous le déguisement du vigilant Batman, va forger une alliance avec un policier nouveau venu à Gotham : le lieutenant James Gordon. Le chef d'oeuvre de Frank Miller et de David Mazucchelli est à nouveau disponible !
Spider-Man se voit offrir un nouveau costume de la part de son nouveau meilleur ami... Tony Stark ! Et les liens tissés aujourd'hui vont déterminer le choix de Spidey durant le conflit à venir. Les Quatre Fantastiques, quant à eux, enquêtent sur un mystérieux objet tombé du ciel tandis que l'équipe la plus secrète de l'univers Marvel, les Illuminati, se reforme. Enfin, les New Warriors deviennent les stars d'une émission de téléréalité et parcourent les routes des États-Unis à la recherche de nouveaux exploits.
