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Le père de l'auteur, Vladek, juif polonais, rescapé d'Auschwitz, raconte sa vie de 1930 à 1944, date de sa déportation. Ce récit est rapporté sous la forme d'une bande dessinée dont les personnages ont une tête d'animal : les juifs sont des souris, les nazis des chats, les Polonais des porcs et les Américains des chiens.
Le Joker, connu pour sa folie, aussi meurtrière que machiavélique, se retrouve face à Batman une nouvelle fois, mais que se passerait-il si l'on inversait les rôles ? Jack Napier, alias le Joker guéri et libéré de ses pensées criminelles se présente aux élections pour devenir le nouveau maire de Gotham. De l'autre côté, le Chevalier Noir tombe dans la criminalité pour sa violence mise à jour par des réseaux sociaux. L'ancien Prince du Crime endosse le rôle du Chevalier Blanc pour sauver Gotham.
Quand le C omédien, justicier au service du gouvernement, se fait défenestrer, son ancien allié, Rorschach, mène l'enquête. I l reprend rapidement contact avec d'autres héros à la retraite dont le D r Manhattan, surhomme qui a modifié le cours de l'histoire. Alors qu'une guerre nucléaire couve entre les USA et l'URSS, tous s'interrogent : qui nous gardera de nos Gardiens ?
